In Memoriam

Stephen T. Warren, PhD, died on June 6, 2021. Warren was born in 1953 and was raised in East Detroit, Mich. He began his studies at Michigan State, graduating with a BS in zoology and a PhD in human genetics, with postgraduate training at the University of Illinois and the European Molecular Biology Laboratory in Heidelberg. Warren was the Charles Howard Candler Chair of Emory’s Department of Human Genetics and the William Patterson Timmie Professor of Human Genetics. He joined the School of Medicine as an assistant professor in biochemistry and pediatrics (medical genetics) and became a full professor in 1993. His research of five decades focused on the mechanistic understanding of fragile X syndrome. In the first major human triumph of the Human Genome Project in 1991, he led an international effort that identified the gene mutation responsible for fragile X. He led clinical trials of drugs to treat the condition. Warren was president of the American Society of Human Genetics, won the William Allan Award, and was an inductee of the National Institute of Child Health and Human Development’s Hall of Honor for the identification of triplet repeat expansion as the cause of fragile X. He was elected to the National Academy of Medicine, the National Academy of Sciences, and the American Academy of Arts and Sciences. Survivors include his wife, Karen and son, Thomas.